Of all the "firsts" on this list, Pakicetus may well be the most counterintuitive. This ultimate whale ancestor, which lived about 50 million years ago, looked like a cross between a dog and a weasel, and walked on four legs just like any other respectable terrestrial mammal. Ironically, the ears of Pakicetus weren't especially well adapted to hearing underwater, so this 50-pound furball probably spent more time on dry land than in lakes or rivers. Pakicetus is also notable as one of the few prehistoric animals ever to be discovered in Pakistan.
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